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Cited article:

A THELYTOKOUS LINEAGE OF SOCIALLY PARASITIC HONEY BEES HAS RETAINED HETEROZYGOSITY DESPITE AT LEAST 10 YEARS OF INBREEDING

Benjamin P. Oldroyd, Michael H. Allsopp, Julianne Lim and Madeleine Beekman
Evolution 65 (3) 860 (2011)
https://doi.org/10.1111/j.1558-5646.2010.01164.x

Pheromonal predisposition to social parasitism in the honeybee Apis mellifera capensis

H.-Q. Zheng, V. Dietemann, R. M. Crewe, et al.
Behavioral Ecology 21 (6) 1221 (2010)
https://doi.org/10.1093/beheco/arq131