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Cited article:

Workers Make the Queens in Melipona Bees: Identification of Geraniol as a Caste Determining Compound from Labial Glands of Nurse Bees

Stefan Jarau, Johan W. van Veen, Robert Twele, et al.
Journal of Chemical Ecology 36 (6) 565 (2010)

Virgin queen execution in the stingless beeMelipona beecheii: The sign stimulus for worker attacks

Stefan Jarau, Johan W. Van Veen, Ingrid Aguilar and Manfred Ayasse
Apidologie 40 (4) 496 (2009)

Some presumably presocial habits of JapaneseCeratina bees, with notes on various social types in Hymenoptera

Shôichi F. Sakagami and Yasuo Maeta
Insectes Sociaux 24 (4) 319 (1977)