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Cited article:

Seasonal variation in the prevalence of Varroa, Nosema and Acarapis in hives from which queen bee mating nuclei are produced

Henry Loeza-Concha, Socorro Salgado-Moreno, Fidel Avila-Ramos, et al.
Journal of Apicultural Research 59 (4) 558 (2020)
https://doi.org/10.1080/00218839.2020.1717060