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Cited article:

Added Weights Lead to Reduced Flight Behavior and Mating Success in Polyandrous Honey Bee Queens (Apis mellifera)

Miranda K. Hayworth, Nels G. Johnson, Matthew E. Wilhelm, Robert P. Gove, Jackie D. Metheny and Olav Rueppell
Ethology 115 (7) 698 (2009)
https://doi.org/10.1111/j.1439-0310.2009.01655.x